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  On-line Guide to Chek Jawa
Phylum Mollusca | Class Bivalvia
 
Bivalves
Class Bivalvia
Fan shell
usually buried in the sand
Bivalves include clams, mussels and oysters. They are abundant on the rocky shore, and buried in the sand and mud.
'Bivalve' means 'two valves'. There are about 7,000 species of bivalves.

Two Shells: A bivalve has a two-part shell, that is, it actually has one shell made up of two parts. Each part of the shell is called a valve. The valves are connected by a hinge and kept shut by one or two large muscles (called adductor muscles). When the bivalve relaxes its adductor muscles, a springy ligament causes the two valves to open. In many bivalves, the hinge between the two valves have teeth to prevent the shells from slipping sideways. Bivalve shells have different shapes and textures to help its owner better survive in its environment.

Drawing by Kelvin LimSanctuary in the Sand: Most bivalves bury themselves. Here they are safer from predators and keep cool and moist during low tide. They use their foot to burrow, then stick out two siphons to the surface. Water is sucked in through one siphon, and ejected through the other.

How do they dig in? A bivalve has only one foot and no other limbs. Yet, it can dig into the sand, some can do it very rapidly indeed. To dig in, the fleshy foot sticks out between the shells. The end of the foot is then expanded into a bulbous shape to form an anchor in the sand or mud. Water is then expelled from between the shells to loosen the sand and mud and the bivalve then quickly contracts its foot to pull itself deeper in. It does this repeatedly until it is at a comfortable depth. Different bivalves bury themselves to different depths. Those with more streamlined shapes dig deeper.

Life in the Slow Lane: Bivalves are mostly sedentary and don't move about as much as most gastropods. Many are adapted to live buried in soft sea bottoms, some live permanently attached to a hard surface. Being mostly immobile, peaceful filter-feeders, most bivalves don't have a head or a radula. Burrowing bivalves have a flattened, blade-like foot to burrow with. Oysters that stick to hard surfaces don't have a foot.

Bivalve Food: Most bivalves use their enlarged mantle cavity to suck water in and to filter out the titbits from the water flow. Cilia (beating hairs) on their gills generate a current through the gills. Mucus on the gills traps food particles which are sent along a groove to the mouth. Fleshy pads near the mouth then push the mucus-food misture into the mouth. The gills also extract oxygen from the water. Oysters and mussels that do not bury themselves simply open their shells a little to get a current going through their bodies. Burying bivalves usually have a pair of siphons, tubes made out of extensions of the mantle. These stick out onto the surface for a one-way flow of water; water enters one siphon and exits the other. In some, these siphons can be quite long so that the bivalve can remain deeply buried and still feed and breathe.

By a Thread: Many bivalves secrete byssal threads, strong protein fibres that can be used to cement themselves to hard surfaces and supports. Burying bivalves may use byssus thread to literally root themselves to the surrounding sand or small stones. The thread is produced by a gland near the foot. The foot gets a grip of the surface and the secretion from the gland flows along a groove in the foot. When the secretion hardens on contact with sea water, the foot is withdrawn.

Bivalve Babies: Most bivalves have separate genders. Bivalves generally practice external fertilisation, releasing their eggs and sperm simultaneously into the water. Most undergo metamorphosis and their larvae look nothing like their adults. Free-swimming larvae develop, drifting with the plankton. These may change as they float, developing a small shell. Eventually, they settle down and develop into miniatures of their parents.

Human uses: Many bivalves are eaten. Although all molluscs can produce pearls, pearls used commercially come mostly from bivalves.

Red Tide: During some seasons, filter feeding bivalves can be highly poisonous. This is because climatic or other conditions favour a bloom of microscopic toxic organisms. These toxins are concentrated in the bivalves. The toxins do not harm the bivalves, but can be fatal to humans who eat the bivalves. These toxic organisms multiply so quickly in ideal conditions that they stain the water red; hence the name 'red tide' for this phenomenon. At other times, filter feeding bivalves may also concentrate other unpleasant chemicals and bacteria which could make you ill.
 

Window-pane shell
lies freely on the sand


Venus clam
buried in the sand


Green mussels
attaches to rocks


Rock oysters
attaches to rocks
 
 
See also ...
Molluscs in general

Links
Bivalvia on the Canada's Aquatic Environments webpage on the University of Guelph website: an easy introduction to the more technical aspects of their morphology, metabolism, reproduction, ecology with an interesting section on their idiosyncracies and photos.
Bivalves on Life on Australian Seashores by Keith Davey on the Marine Education Society of Australia website: a fabulous goldmine of a site which covers various kinds of intertidal habitats and their inhabitants. Fact sheet on bivalves and details on an oyster found in Australia.
Class Bivalvia on Biomedia of the Glasgow University Zoological Museum on the Biological Sciences, University of Paisley, Scotland website: a brief introduction with explanations of the major classes, a glossary of terms and diagrams and photos.
Bivalves on The Gladys Archerd Shell Collection at Washington State University Tri-Cities Natural History Museum website: brief fact sheet on bivalves with links to various species.

Other references
  • Barnes, Robert D. & Ruppert, Edward E., 1996. Invertebrate Zoology. Harcourt College Publishers. 6th Edition. pp. 1056, G-1-16, I-1-30.
  • Pechenik, Jan A., 2000. Biology of the Invertebrates. McGraw-Hill Book Co., Singapore. 578 pp.
  • Hendler, Gordon, John E. Miller, David L. Pawson and Porter M. Kier, 1995. Sea Stars, Sea Urchins, and Allies: Echinoderms of Florida and the Caribbean. Smithsonian Institution Press. 390 pp.
  • Schoppe, S., 2000. Echinoderms of the Philippines. Times Edition, Singapore. 144 pp.
  • Tan, Leo W. H. & Ng, Peter K. L., 1988. A Guide to Seashore Life. The Singapore Science Centre, Singapore. 160 pp.

 

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